- Received August 23, 2016.
- Accepted November 17, 2016.
- Published online December 14, 2016.
Copyright & Usage
© 2016 The Authors. Published by the Royal Society under the terms of the Creative Commons Attribution License http://creativecommons.org/licenses/by/4.0/, which permits unrestricted use, provided the original author and source are credited.
- 1Department of Zoology, University of Cambridge, Cambridge, UK
- 2Institute of Evolutionary Biology, University of Edinburgh, Edinburgh, UK
- 3Division of Evolution, Ecology and Genetics, Research School of Biology, The Australian National University, Canberra, Australia